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Subtle behavioral variation in wild chimpanzees, with special reference to Imanishi's concept of kaluchua
  Nakamura, Michio, Nishida, Toshisada (2006-01)
  Primates, 47(1): 35-42
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Ontogeny of a social custom in wild chimpanzees: age changes in grooming hand-clasp at Mahale
  Nakamura, Michio, Nishida, Toshisada (2013-02)
  American journal of primatology, 75(2): 186-196
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